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I stumbled upon this paper sometime ago and I think it could be really useful in implementing (mollusk-like) shell design in the game:
David M. Raup. (1966). Geometric Analysis of Shell Coiling: General Problems. Journal of Paleontology, 40(5), 1178–1190. doi:10.2307/1301992
I already posted it in the community forums, see if you like it
I think another major extinction event could be major solar flares which would strip away your planets atmosphere, making the UV rays much bigger and needing you to use photosynthesis things to up your UV resistance to about 105 or 110% to survive, while also heavily increasing the tempearture, and all this would last only 1 generation since atmospheres heal quick iirc
Like I wrote in another thread, more major events can be the movement of the tectonic plates, major continental collisions or rifting or the re-activation of hotspots even, other than classic events such as the meteorites or eruptions
Well, this thread is for events like this in general so lets move on from algal blooms and suggest something else, like antimatter creatures? (jk)